It is difficult to know exactly when the shift from gelatine to agar occurred. As often happens for scientific breakthroughs, agar was likely adopted incrementally alongside the use of other growth media. In 1913, for example, the first diagnosis of Serratia marcescens as a human pathogen was made by growing it on agar as well as on potatoes.
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The answer is essentially hardware-level dependency injection. Before calling LD_DESCRIPTOR, the caller saves its desired test constant into a hardware latch using a micro-op called PTSAV (Protection Save). Within LD_DESCRIPTOR, another micro-op called PTOVRR (Protection Override) retrieves and fires the saved test.